Pharmacology students in matching vests


The Pharmacology Training Program currently has 31 students enrolled in the Ph.D. Program. Of the current students, 5 entered the program as M.D./Ph.D. (MSTP) students; and 4 others entered the program from the umbrella Biomedical Science Program (BSP). The Pharmacology Graduate Training Program consistently matriculates students from all three potential sources: direct recruiting, the BSP, and the MSTP. With an average GPA of 3.6 and GRE scores of V: 157, Q: 157(170 scale), our applicants are clearly competitive. They also represent a broad and diverse geographic distribution as well as a variety of undergraduate majors including: biology, molecular biology, biochemistry, chemistry, physics, pharmacy and psychology. Over the last ten years, the average size of the matriculated class of students Pharmacology Training Program students has been 4.6 and the average time to degree is ~5.2 years. The retention rate, that is students who complete the program with their Ph.D. is (37/49) or 76% over the last 10 years. Another 8 students completed the program with M.S. degrees. Thus, the overall percentage of a students graduating with an advanced degree is (45/49) 92%. Between 1998 and 2017, the Pharmacology T32 Training Grant supported 65 of its 111 Training Program graduate students.

Current StudentsStudent Information
Faith AguilarFaith Aguilar
BS, Chemistry, Univ. of Texas Rio Grande Valley, Edinburg, TX, 2017
Lorena Alamillo

Lorena Alamillo
BS, Biochemistry, Metropolitan State University of Denver, Denver, CO, 2019


Dillon BoultonDillon ​Boulton
BS, Biology, Rowan Univ., Glassboro, NJ, 2016
Mentor: Caino, M. Cecilia
Elucidating the role of Mitochondrial Rho-like GTPase, Miro2, in the regulation of mitochondrial dynamics and turnover
Nicole Brown

Nicole Brown
BA, Neuroscience, Univ. of Colorado, Boulder, CO, 2016
Mentor: Bayer, Ulli
I study how LTP vs LTD crosstalk occurs between subunits within the CaMKII 12-meric holoenzyme
Nicole Brown's Publications on NIH

Amelia BurchAmelia Burch
BS, Neuroscience, Emory Univ., Atlanta, GA, 2015
Mentor: Herson, Paco
The role of the TRPM2 ion channel in synaptic and cognitive dysfunction following focal cerebral ischemia
Amelia Burch's Publications on NIH
Nan ChenNan Chen
BS, Pharmacy, China Pharmaceutical Univ., 2014; M.S., Drug Discovery and Development, Drexel Univ., 2016
Mentor: Doebele, Robert C.
Investigating how EGFR mediates early adaptive resistance to tyrosine kinase inhibitors in NSCLC driven by oncogenic fusion kinases
Nan Chen's Publications on PubMed
Lily FeldmanLily Elizabeth Feldman
BS, Biochemistry, University of New Mexico, Albuquerque, NM, 2020
Mentor: Costello, C. James
Characterizing novel chemoresistance mechanisms in human bladder cancer
Michelle Foreman

Michelle Foreman
BS, Mathematical Biology, University of Houston, Houston, TX, 2019
Mentor: Black, Joshua
Investigating epigenetic dysregulation of the interferon receptor locus by PRC1 in trisomy 21

Michelle's content on RNA Journal
Michelle's publications on PubMed

Madison FurnishMadison Furnish
BS, Biology, Univ. of Louisville, Louisville, KY, 2017
Mentor: Caino, M. Cecilia
Identification and characterization of novel effector proteins in the Rho-like GTPase Miro2 that alter metabolism and motility in prostate cancer.
Madison's publications on PubMed
Joshua GarciaJoshua Garcia
BS, Biology, Univ. of New Mexico, 2015
Mentor: Smith, Katharine R.
Understanding mechanisms of inhibitory synapse down-regulation in vulnerable cell-types during OGD
Sean HickeySean Hickey
BA&Sc., Psychology and Philosophy, Regis Univ., Denver, CO, 2013
Mentor: Saba, Laura M.
Elucidating genetically regulated gene expression in the context of addiction.
Sean Hickey's Publications on Research Gate
Jessica HsuJessica Hsu
BS, Molecular Toxicology, Univ. of California Berkeley, Berkeley, CA, 2014
Mentor: Ford, Heide L.
Utilizing zebrafish as a model organism to investigate the mechanisms by which developmental genes contribute to formation and progression of Rhabdomyosarcoma.
Alex Huffman

Alex Huffman
BA, Neuroscience, Northwestern Univ., Evanston, IL, 2018
Mentor: Dell’Acqua, Mark
Investigating the signaling of AKAP-anchored PKA in synaptic plasticity.

Connor HughesConnor Hughes
BS, Biology, Univ. of Richmond, Richmond, VA, 2016
Mentor: Ford, Heide L.
I study the role of Eya transcriptional cofactors in regulation of the immune response in triple negative breast cancer
Danielle Jeffrey

Danielle Jeffrey
BS, Cellular and Molecular Biology, California State Univ., Chico, CA, 2012; M.S., Biomedical Sciences, Regis Univ., Denver, CO, 2016
Mentor: Dabertrand, Fabrice
Investigating the role of mural cells in normal brain microcirculation in comparison to small vessel diseases.

Austin JollyAustin Jolly
BA, Intergrated Physiology, Univ. of Colorado, Boulder, CO, 2016
Mentor: Weiser-Evans, Mary C.M.
Investigating the contribution of vascular progenitor cells to pathologic vascular remodeling, fibrosis, and hypertension.
Dean KareemoDean Kareemo
BS, Pharmaceutical Sciences, Univ. of Rhode Island, Kingston, RI, 2018
Mentor: Kennedy, Matt
Investigating the unique trafficking pathways of synaptic proteins

Mason KleinjanMason Kleinjan
BS, Biology, Univ. North Dakota, Grand Forks, ND, 2017
Mentor: Kennedy, Matthew J.
Investigating the Spine Apparatus and unconventional protein trafficking in neurons
Christophe Langouet-AstrieChristophe Langouet-Astrie
BS, Biotechnology & Statistics Minor, James Madison Univ., Harrisonburg, VA, 2015; M.S., Biomedical Sciences and Biotechnology, Univ. of Colorado Anschutz, Denver, CO, 2018
Mentor: Schmidt, Eric P.
Glycosaminoglycan therapeutics to sequester pathogenic toxins in Sepsis.
Christophe Langouet-Astrie's Publications on PubMed
Jiajun LiJiajun Li
BS, Biological Science, Ocean University of China, 2015; MS, Biology, Illinois Institute of Technology, Chicago, IL, 2017
Mentor: Lahm, Tim
Estrogen signaling effects on Right Ventricle during adaptive and maladaptive phases of Pulmonary Hypertension.
Jiajun Li's Publications on NIH
Brian LloydBrian Lloyd
BM, Vocal Performance, Univ. of Denver, Denver, CO, 2013; BS, Biology, Univ. of Colorado, Denver, CO, 2017
Mentor: Aoto, Jason
Subsynaptic characterization of neurexins
Brian Lloyd's Publications on PubMed
Tyler MartinezTyler Martinez
BA, Physics & French, Vassar College, Poughkeepsie, NY, 2011
Mentor: Dell'Acqua, Mark L.
Synapse regulation by NFAT-dependent transcription.
Carley MillerCarley Miller
BS, Neuroscience, Penn State University, State College, PA, 2018
Mentors: Heinsbroek, Jasper A. & Aoto, Jason
Characterizing the role of the ventral pallidum in stress and drug use disorders
Carley Miller's Publications on PubMed
orman headshot

Michael Orman
BS Chemical & Biomolecular Engineering, Georgia Institute of Technology, Atlanta, GA, 2015
MS Biotechnology, Northwestern Univ., Evanston, IL, 2018

Mentor: Costello, Jim
Investigating molecular drivers of metastatic prostate cancer

Madison RodriguezMadison Rodriguez
BA, Biochemistry, University of Colorado Boulder, Boulder, CO, 2019
Ian SheltonIan Shelton
BA, Science and Management, Claremont McKenna College, Claremont, CA, 2017
Mentor: Jordan, T. Craig
Characterizing the role of leukemia stem cells to better understand the underlying cellular and molecular mechanisms driving drug resistance in Acute Myeloid Leukemia
Emilia SmithEmelia Smith
BS, Molecular and Cellular Biology, Univ. of Illinois at Urbana-Champaign, IL, 2019
Mentor: Ford, Heide
Investigating molecular crosstalk mechanisms between phenotypic clones within the heterogeneous tumor and how to target this intratumoral signaling to prevent drug resistance and progression towards metastatic disease
stokesEric Stokes
BS, Biology, Morehouse College, Atlanta, GA, 2018
Mentor: Aoto, Jason

Investigating Nrxn3a influence on synapse organization and transmission

Alexander Strait​Alexander (Sascha) Strait
BS, Biological Science, Univ. of California, Davis, CA, 2004
Mentor: Wang, Xiao-Jing
TGF-β signaling regulates the response to immune checkpoint blockade in head and neck squamous cell carcinoma.
Alexandra Strait's Publications on PubMed
Philip TatmanPhilip Tatman
BS, Physiology and Biochemistry, Univ. of Washington, Seattle, WA, 2014
Mentor: Black, Joshua C.
Activation of the insulin receptor by histone three tail peptides.
Publications: View on Google Scholar
Amber Truitt
Amber Truitt
BS, Chemistry, Towson University, MD, 2014
Jonathan TullisJonathan Tullis
BS, Chemistry, Santa Clara Univ., Santa Clara, CA, 2014
Mentor: Bayer, K.​​​​​​​​ Ulrich
Investigating DAPK1 localization and function in synaptic plasticity.
Jonathan Tullis's Publications on NIH
Jackie Turner

Jaqueline Turner
BA, Biochemistry and Integrative Physiology, Univ of Colorado Boulder., Boulder, CO, 2017
Torres, Raul M. 
Lipid modulation of anti-tumor immunity and immunosurveillance

Jackie Turner's Publications on NIH

Logan TylerLogan Tyler
BS, Biology, Villanova Univ., 2012; MPH., Epidemiology, Drexel Univ., 2014
Mentor: Doebele, Robert C.
Investigating mechanisms of acquired resistance to tyrosine kinase inhibitors (TKIs) in human primary cell lines with driving receptor tyrosine kinase (RTK) fusion mutations
Logan Tyler's Publications on PubMed
Gregory WrightGregory Wright
BS, Chemistry, Rochester Institute of Technology, Rochester, NY, 2003; MS Chemistry & Bioinformatics, Rochester Institute of Technology, Rochester, NY, 2014
Mentor: Black, Joshua C.
Investigating transient site-specific gain as a mechanism of gene amplification
Nathaniel XanderNathaniel Xander 
BS, Biochemistry, Temple University, Philadelphia, PA, 2017; Prof Science Masters, Biotechnology, Temple University, Philadelphia, PA, 2019